Joe Lo Truglio doesn’t have many acting credits to his name, but his distinctive appearence and frequent collaboration with Judd Apatow have made him into a memorable “That Guy” of comedy. Lo Truglio attended New York University, where he participated in the school’s sketch comedy group. It was there that he met a group of people (including David Wain, Michael Ian Black, Thomas Lennon and Ben Garant) who would go on to form the cast of the MTV sketch comedy show The State. After The State ended in 1995, Lo Truglio continued to act in television, appearing in shows such as Upright Citizens Brigade, Law & Order, Third Watch, and Reno 911!. In 2001, Lo Truglio reunited with David Wain to work on Wet Hot American Summer, which gave him a bit of a career boost, but it wasn’t until Superbad that I really took notice of him as a skeezy driver who escorts Michael Cera and Jonah Hill to a house party where he isn’t welcome. This role catapulted Lo Truglio into several other big-screen comedies such as I Love You, Man, Fanboys, Paul, and Pineapple Express. But I think Lo Truglio’s best work to date was his role in Role Models as a hardcore LARPer who gets me laughing every time he’s on screen. Lo Truglio turns 41 today.